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Movement of large bluefin tuna, Thunnus thynnus, in the North Pacific Ocean, as determined from the Japanese longline fishery, and implications regarding interactions between the fisheries of the western and eastern Pacific Ocean

Patrick K. Tomlinson
Inter-American Tropical Tuna Commission
c/o Scripps Institution of Oceanography
8604 La Jolla Shores Drive
La Jolla, California 92037 USA

ABSTRACT

Japanese high-seas longline fishery data were used to plot the monthly distribution of catch and effort by 5-degree area. A scheme of weighting each 5-degree square was devised so that the location of catch or effort could be easily determined. The results show that there is a very pronounced pattern of movement in both the locations of the catch and the locations of the effort. Since the effort was more broadly distributed than the catches, it was concluded that large bluefin of the size caught by longliners generally follow a pattern of moving to the west in the spring and to the east in the fall, with most of the fish of this size staying in the northern Pacific Ocean. Occasionally, the eastern extension of the movement makes these large bluefin available to the purse seine fishery in the eastern Pacific Ocean.

1. INTRODUCTION

This study is based on catch and effort data for the Japanese high-seas longline fishery kindly made available to the staff of the Inter-American Tropical Tuna Commission (IATTC) by the National Research Institute of Far Seas Fisheries of Japan.

2. METHODS

The data utilized include summaries of catch by species, in numbers of fish, by year (1952-1986), month, and 5-degree area, and summaries of the effort, in number of hooks (not adjusted for any changes in efficiency over time), by year, month, and 5-degree area. The data for northern bluefin tuna (NBF) were extracted and summarized by month and 5-degree area. For the purpose of showing the distribution of catch by month, two time periods of 14 years each--1959 to 1972 and 1973 to 1986 were selected. Maps of the distributions of catches for each of the two time periods (Figures 1 to 12, top panel for the early period and bottom for the late period) were prepared. After examination of these distributions, a NBF core area was chosen (Figure 13). The core area contains 58 5-degree areas, all in the northern hemisphere. Let

Cijk = NBF catch for year i, month j, and 5-degree area k,

Eijk = number of hooks for year i, month j, and 5-degree area k,

and

Uijk = catch per hook (CPUE) for year i, month j, and 5-degree area k,

Uijk = Cijk/Eijk

where

i = 1, 2, ...., N; N = number of years,
j = 1, 2, ...., M; M = number of months, and
k = 1, 2, ...., L; L = number of 5-degree areas.

Total catches by year for the entire Pacific Ocean (SSS Cijk, N = 1, M = 12, L = 636) and for the core area (SSS Cijk, N = 1, M = 12, L = 58) were compiled (Table 1). The monthly totals, by year, of NBF catch, in numbers of fish, were calculated for the entire Pacific Ocean (SSS Cijk, N = 1, M = 1, L = 636) (Table 2) and for the core area (SSS Cijk, N = 1, M = 1, L = 58) (Table 3). The monthly totals, by year, of longline effort in thousands of hooks, were calculated for the core area (SSS Eijk, N = 1, M = 1, L = 58) (Table 4).

Within each year, a search was made to find the 5-degree area-month strata within the core area which contained the greatest catches and the greatest effort (Table 5). Similarly, the numbers of 5-degree area-month strata with catches of at least one NBF (Table 6), and the numbers of 5-degree area-month strata with effort of at least one hook (Table 7) were counted for each year.

For each longitudinal strip of 5 degrees, starting at the equator and going north or south, each 5-degree area going from 80°W to 80°E was given a number from 1 to 40. For each strip, the numbering starts with 1 in the east and goes sequentially to 40 in the west. The area covered by this numbering system has a total of 880 5-degree areas (22 5-degree strips with 40 5-degree areas per strip), with half the areas north of the equator. An index is formed by multiplying the number of NBF caught in a 5-degree area-month by the numerical value of that area and then summing over all area-months. This sum is divided by the sum of the catches to obtain a weighted estimate of the average index. Let

Xk = k = east-west value of area k; k = 1, 2, ...., 40, and

Vij = SS XkCijkl/SS Cijkl; l = 1, 2, ...., 22 = east-west value for year i and month j.

This provides a numerical representation of the east-west origin of the catch; the higher the value, the more westerly the catch. These east-west values were computed for each month of each year of the 1952-1986 period for the entire Pacific Ocean (Figure 14) and for the NBF core area (Figure 15). Similar east-west values were calculated for effort within the entire Pacific Ocean (Figure 16), effort within the core area (Figure 17), and CPUE within the core area (Figure 18). The monthly values of catch (dotted lines) and effort (solid lines) for each year were plotted for the entire Pacific Ocean (Figure 19) and for the NBF core area (Figure 20) to help determine if the east-west location of catch was influenced by the east-west location of effort.

3. RESULTS

3.1 Longline Catches

The core area usually contained more than 85% of the catch during the 1952-1971 period, but after that the percentages became more variable (Table 1). The data of Nakamura (1969) and Shingu et al. (1974) give the impression that most NBF caught by longline are between about 80 and 200 kg. The longline fishery appears to have made important contributions to the total catch during the early 1960s, but since about 1965 the contribution has been minor.

3.2 Movement

It appears that the large NBF caught by longliners concentrate on or near the spawning grounds in April through June and then move slightly to the north and substantially to the east (Figures 1 to 12, top and bottom, and Figure 17). The fish reach the central Pacific Ocean and beyond, especially during December through February. The distance between the average western location (say 25°N-130°E) in May and the average eastern location (say 32°N-172°E) in December-January is about 2.3 thousand nm (4.2 km). This means that the large NBF move about 383 nm (700 km) per month, or about 13 nm (23 km) per day. Of course, there is considerable variability in the data.

The monthly variation in the east-west location of the catch for the entire Pacific Ocean shows a consistent pattern (Figure 14), while the variation for Pacific-wide effort (Figure 16) is not so consistent. The effort is distributed farther to the east than the NBF catch for the Pacific-wide data, and the timing of the movements for catch (Figure 19, dotted line) differ from those of effort (Figure 19, solid line). However, the results are somewhat different when the analysis is confined to the core area. For the core area, the oscillation for catch (Figure 15) and effort (Figure 17) are much more nearly similar, with similar timing for catch (Figure 20, dotted line) as for effort (Figure 20, solid line). The locations of the greatest CPUEs (Figure 18) within the core area are very similar to the locations of the greatest catches (Figure 15). If the monthly indices for CPUEs are regressed against the indices for catch, the regression explains 89 percent of the variation in the CPUE data and the slope is not significantly different from 1. However, if a similar regression is done with effort, it explains only 51 percent of the variation in the effort data, and the slope is significantly different from 1. In the core area (on a month-by-month basis) the number of 5-degree areas with effort averaged between 11 and 51 (Table 7) with the overall average at about 30, while the number of 5-degree areas with catch averaged between 0 and 24 (Table 6) with the overall average at about 10. Therefore, within the core area, about two-thirds of the month-5 degree areas with effort produced no catch of NBF.

4. DISCUSSION

It is unfortunate that the results on movements must be tempered by the limitations of the data. Most of the NBF are caught from the core area, and within this area the effort moves west to east and then back to the west on a regular cycle which is similar to that of the catch location, but more spread out. Since the distribution of effort is more spread out than the distribution of catch, there must be a high correlation between where the NBF are caught and where they actually are located. Also, the greatest CPUEs for NBF are in the same areas and times as the greatest catches. In other words, the effort seems to bracket the area where the NBF are being caught, with areas surrounding the catch areas having zero CPUE. Figures 21 and 22, which show the locations of catches and locations of areas with effort, but no catch, for two different months (May and December), clearly indicate that effort extends beyond the range of the catch, especially when the catches are from offshore during December. When the areas with catches were within the southwest part of the core area, there were still some effort to the east of where the NBF were being caught, but not much. However, when the main areas with catches were in the northeast part of the core area, there were still large amounts of effort in the western part with small catches of NBF.

Large NBF, similar in size to those caught by longliners, were caught by purse seiners in the eastern Pacific during November and December of 1988 (Foreman and Ishizuka, 1990). These fish may have appeared in the eastern Pacific during the fall of 1988 because they were distributed further to the east than is usually the case. It does indicate there is some possibility that the large NBF usually exploited by longliners in the central Pacific during December to February could be taken by purse seiners operating in the eastern Pacific.

5. LITERATURE CITED

Foreman, T.J., and Y. Ishizuka. 1990. Giant bluefin off Southern California, with a new California size record. Calif. Fish Game 76(3): 181-186.

Nakamura, H. 1969. Tuna Distribution and Migration. London, Fishing News (Books) Ltd. 76 p.

Shingu, C., Y. Warashina, and N. Matsuzaki. 1974. Distribution of bluefin tuna exploited by longline fishery in the western Pacific Ocean. Bull. Far Seas Fish. Res. Lab. 10: 109-140.

Figure 1. January distribution of Japanese longline catches of NBF, summed over two time periods of 14 years each (1959 to 1972).

Figure 1. January distribution of Japanese longline catches of NBF, summed over two time periods of 14 years each (1973 to 1986).

Figure 2. February distribution of Japanese longline catches of NBF, summed over two time periods of 14 years each (1959 to 1972).

Figure 2. February distribution of Japanese longline catches of NBF, summed over two time periods of 14 years each (1973 to 1986).

Figure 3. March distribution of Japanese longline catches of NBF, summed over two time periods of 14 years each (1959 to 1972).

Figure 3. March distribution of Japanese longline catches of NBF, summed over two time periods of 14 years each (1973 to 1986).

Figure 4. April distribution of Japanese longline catches of NBF, summed over two time periods of 14 years each (1959 to 1972).

Figure 4. April distribution of Japanese longline catches of NBF, summed over two time periods of 14 years each (1973 to 1986).

Figure 5. May distribution of Japanese longline catches of NBF, summed over two time periods of 14 years each (1959 to 1972).

Figure 5. May distribution of Japanese longline catches of NBF, summed over two time periods of 14 years each (1973 to 1986).

Figure 6. June distribution of Japanese longline catches of NBF, summed over two time periods of 14 years each (1959 to 1972).

Figure 6. June distribution of Japanese longline catches of NBF, summed over two time periods of 14 years each (1973 to 1986).

Figure 7. July distribution of Japanese longline catches of NBF, summed over two time periods of 14 years each (1959 to 1972).

Figure 7. July distribution of Japanese longline catches of NBF, summed over two time periods of 14 years each (1973 to 1986).

Figure 8. August distribution of Japanese longline catches of NBF, summed over two time periods of 14 years each (1959 to 1972).

Figure 8. August distribution of Japanese longline catches of NBF, summed over two time periods of 14 years each (1973 to 1986).

Figure 9. September distribution of Japanese longline catches of NBF, summed over two time periods of 14 years each (1959 to 1972).

Figure 9. September distribution of Japanese longline catches of NBF, summed over two time periods of 14 years each (1973 to 1986).

Figure 10. October distribution of Japanese longline catches of NBF, summed over two time periods of 14 years each (1959 to 1972).

Figure 10. October distribution of Japanese longline catches of NBF, summed over two time periods of 14 years each (1973 to 1986).

Figure 11. November distribution of Japanese longline catches of NBF, summed over two time periods of 14 years each (1959 to 1972).

Figure 11. November distribution of Japanese longline catches of NBF, summed over two time periods of 14 years each (1973 to 1986).

Figure 12. December distribution of Japanese longline catches of NBF, summed over two time periods of 14 years each (1959 to 1972).

Figure 12. December distribution of Japanese longline catches of NBF, summed over two time periods of 14 years each (1973 to 1986).

Figure 13. Japanese longline fishery core area for NBF. Total area contains 58 5-degree areas.

Figure 14. Japanese longline monthly index of east-west location for NBF catches from the entire Pacific Ocean. The greater the number, the farther west the NBF were captured, on average.

Figure 15. Japanese longline monthly index of east-west location for NBF catches from the core area. The greater the number, the farther west the NBF were captured, on average.

Figure 16. Japanese longline monthly index of east-west location for effort from within the entire Pacific Ocean. The greater the number, the farther west the effort, on average.

Figure 17. Japanese longline monthly index of east-west location for effort from within the core area. The greater the number, the farther west the effort, on average.

Figure 18. Japanese longline monthly index of east-west location for CPUEs from within the core area. The greater the number, the farther west the greatest CPUE, on average.

Figure 19. Annual comparisons of the Japanese longline monthly index of east-west location for the NBF catches (dotted line) from the entire Pacific Ocean to those of the effort (solid line). (A)

Figure 19. Annual comparisons of the Japanese longline monthly index of east-west location for the NBF catches (dotted line) from the entire Pacific Ocean to those of the effort (solid line). (B)

Figure 19. Annual comparisons of the Japanese longline monthly index of east-west location for the NBF catches (dotted line) from the entire Pacific Ocean to those of the effort (solid line). (C)

Figure 20. Annual comparisons of the Japanese longline monthly index of east-west location for the NBF catches (dotted line) from the core area to those of the effort (solid line). (A)

Figure 20. Annual comparisons of the Japanese longline monthly index of east-west location for the NBF catches (dotted line) from the core area to those of the effort (solid line). (B)

Figure 20. Annual comparisons of the Japanese longline monthly index of east-west location for the NBF catches (dotted line) from the core area to those of the effort (solid line). (C)

Figure 21. Annual distributions of Japanese longline catches of NBF taken within the core area during May. Catches are indicated by *, +, o, or - (with the greatest to least catches in the same order). Areas with effort, but no catch, are indicated by dots.

Figure 21 a.

Figure 21 b.

Figure 21 c.

Figure 22. Annual distributions of Japanese longline catches of NBF taken within the core area during December. Catches are indicated by *, +, o, or - (with the greatest to least catches in the same order). Areas with effort, but no catch, are indicated by dots.

Figure 22 a.

Figure 22 b.

Figure 22 c.

Table 1. Japanese longline catches in number of northern bluefin tuna from the core area and the entire Pacific Ocean.

YEAR

CORE CATCH

ENTIRE CATCH

DIFF.
ENTIRE - CORE

% DIFF. =
100*DIFF/ENTIRE

52

23853

24012

159

0.662

53

22098

22336

238

1.066

54

24125

24472

347

1.418

55

23064

23268

204

0.877

56

21293

22845

1552

6.794

57

10598

11006

408

3.707

58

6480

6684

204

3.052

59

27741

32604

4863

14.915

60

50599

51926

1327

2.556

61

50755

53833

3078

5.718

62

42970

45715

2745

6.005

63

40690

44222

3532

7.987

64

19755

20706

951

4.593

65

15342

17314

1972

11.390

66

7432

9006

1574

17.477

67

5788

6216

428

6.885

68

3568

3756

188

5.005

69

2588

2862

274

9.574

70

1628

1826

198

10.843

71

1733

2030

297

14.631

72

615

801

186

23.221

73

596

1030

434

42.136

74

583

892

309

34.641

75

342

1440

1098

76.250

76

478

1149

671

58.399

77

2105

2275

170

7.473

78

3034

3099

65

2.097

79

1958

2237

279

12.472

80

871

1216

345

28.372

81

2336

2576

240

9.317

82

2290

2437

147

6.032

83

776

882

106

12.018

84

570

748

178

23.797

85

324

412

88

21.359

86

295

395

100

25.316


Table 2. Japanese longline catches of northern bluefin tuna in numbers of fish from the Pacific Ocean.

YEAR

JAN.

FEB.

MAR.

APR.

MAY

JUNE

JULY

AUG.

SEP.

OCT.

NOV.

DEC.

52

1211

277

105

613

7865

1961

2280

6224

2188

238

250

800

53

640

602

70

678

8818

5039

866

1808

2117

1061

439

198

54

760

411

554

1127

8144

2533

3759

3250

2729

840

273

92

55

351

537

408

2116

5724

1572

298

5660

3833

2123

510

136

56

235

184

437

2470

3662

1069

546

9437

3648

635

253

269

57

227

331

482

1886

2679

601

1181

2592

294

36

87

610

58

273

294

459

1023

765

461

17

101

1014

648

873

756

59

1014

1255

1183

4960

3707

3673

2021

3715

2967

1528

2845

3736

60

1061

704

3554

17384

10115

7119

873

1427

2781

1665

4532

711

61

1136

1306

3844

10297

15398

4369

3752

3433

4983

1514

1784

2017

62

1668

1355

2672

10719

16791

5206

669

1075

2057

627

1009

1867

63

884

382

2100

14799

18342

1949

335

2237

1497

589

425

683

64

581

176

1066

5603

7672

1676

227

772

1251

483

685

514

65

258

366

1369

3356

5988

2901

175

276

532

610

888

595

66

91

134

439

1432

2269

1083

593

397

866

940

367

395

67

387

226

548

871

1463

406

148

592

987

247

187

154

68

119

321

752

398

589

322

168

254

367

179

146

141

69

183

172

244

223

526

174

152

274

396

204

103

211

70

99

128

112

201

438

316

107

29

60

45

215

76

71

251

438

449

54

265

197

30

22

53

70

48

153

72

160

73

46

58

86

89

26

30

33

14

80

106

73

160

73

119

61

72

265

36

44

40

43

45

72

74

35

21

38

40

116

170

71

102

150

45

65

39

75

120

34

28

33

91

121

32

200

438

295

19

29

76

224

56

87

80

186

118

47

220

33

19

20

59

77

155

460

453

722

131

76

23

3

178

36

2

36

78

71

246

526

203

101

43

15

175

1607

35

13

64

79

123

130

241

304

98

182

63

172

720

99

48

57

80

143

51

109

179

251

82

54

50

29

25

80

163

81

373

412

677

274

81

62

26

247

133

63

130

98

82

151

293

193

162

206

424

740

87

40

37

39

65

83

99

121

219

145

46

90

16

28

16

17

33

52

84

175

133

180

10

32

17

44

54

37

16

16

34

85

84

78

74

23

9

27

28

18

13

18

20

20

86

83

70

56

15

8

66

31

18

12

10

13

13

AVG

388

339

683

2358

3507

1270

556

1286

1089

430

473

429


Table 3. Japanese longline catches of northern bluefin tuna in numbers of fish from the core area.

YEAR

JAN.

FEB.

MAR.

APR.

MAY

JUNE

JULY

AUG.

SEP.

OCT.

NOV.

DEC.

52

1211

277

105

598

7839

1941

2226

6195

2183

238

250

790

53

620

602

67

669

8785

5019

832

1757

2066

1044

439

198

54

660

383

550

1075

8126

2511

3704

3238

2718

821

262

77

55

342

534

408

2075

5695

1565

261

5628

3805

2111

504

136

56

233

184

427

2398

3655

1040

517

8868

3078

569

57

267

57

225

331

431

1837

2661

574

1164

2374

285

28

80

608

58

261

292

457

1015

760

439

4

20

975

640

865

752

59

811

1225

1179

4938

3645

3651

829

533

2930

1446

2818

3736

60

1061

689

3552

17366

10092

7102

382

1015

2542

1628

4461

709

61

1131

987

3827

10201

15378

3951

3604

2655

3725

1506

1778

2012

62

1625

1148

2594

10462

16682

5092

414

599

985

585

1000

1784

63

831

346

2041

14698

18308

1629

101

140

1189

381

369

657

64

576

176

1020

5484

7624

1634

104

422

1131

423

669

492

65

170

269

1327

3209

5591

2702

52

46

400

539

444

593

66

72

134

425

1060

2151

835

120

192

808

898

362

375

67

386

222

546

833

1389

331

16

525

969

245

183

143

68

119

287

749

385

570

278

126

236

361

173

144

140

69

181

166

232

188

472

156

107

231

364

179

103

209

70

96

124

81

197

424

308

17

15

54

34

204

74

71

240

371

436

52

232

129

2

2

28

51

47

143

72

136

39

46

40

73

58

7

2

27

11

77

99

73

143

56

118

51

46

37

4

6

16

14

42

63

74

30

20

31

22

77

62

38

80

125

20

46

32

75

120

19

20

23

37

16

7

21

32

8

14

25

76

45

24

44

35

35

7

5

184

5

18

19

57

77

133

441

447

716

103

16

3

2

173

33

2

36

78

68

242

522

199

99

28

6

161

1600

33

12

64

79

115

89

238

287

70

130

1

145

687

96

44

56

80

143

35

102

146

106

58

30

0

10

12

79

150

81

362

360

655

258

44

39

3

222

110

59

128

96

82

148

286

184

159

188

413

721

20

35

37

39

60

83

81

94

214

140

41

82

7

16

8

17

28

48

84

170

117

178

10

5

9

4

10

6

16

16

29

85

73

73

71

22

7

11

3

2

6

18

19

19

86

80

67

55

12

3

27

0

4

12

9

13

13

AVG

363

306

668

2310

3458

1197

441

1016

956

398

446

421


Table 4. Japanese longline fishing effort, in thousands of hooks, in the core area.

YEAR

JAN.

FEB.

MAR.

APR.

MAY

JUNE

JULY

AUG.

SEP.

OCT.

NOV.

DEC.

52

17253

13048

6278

5065

6390

1607

2354

4338

2681

4009

6686

9060

53

15815

20780

4544

2264

7190

3995

1873

1897

2074

4361

6893

12059

54

14640

19319

17421

7914

8230

5848

2850

2266

2605

3771

4925

8140

55

14468

16626

15809

10539

9133

3098

364

2127

3747

4723

8870

11818

56

13963

15107

16367

9945

8129

4486

348

739

2796

5591

9827

9018

57

14497

16868

14142

8181

6146

3110

355

1141

2413

3619

7521

13621

58

18609

14059

11809

8568

6499

3469

450

1113

3561

7555

12365

11206

59

13620

13455

14129

6412

5823

4463

3299

2967

9492

8874

12051

12183

60

17197

17919

15242

9580

8534

5906

3358

2449

9096

12078

12803

15312

61

17312

15643

16157

11075

9133

6099

6462

6121

5142

7451

11638

12250

62

13151

13348

14524

8264

7083

3216

863

1627

5149

5862

8898

8156

63

8939

9824

11629

10569

7089

1371

565

1033

4024

7620

10594

15655

64

10063

8978

9974

6696

5871

2787

1375

2917

5787

7571

5341

5140

65

8572

9381

9062

6120

5591

3337

2895

5035

7845

8808

8543

8050

66

8300

10694

10679

5917

4319

2702

3059

4973

7964

10485

9983

9918

67

14449

14944

16990

7504

4555

2259

1476

5368

9323

13524

14733

12702

68

14168

13149

11666

4626

2892

2255

2436

3017

6613

10351

13233

12156

69

15191

13415

11167

3470

2261

2162

2739

2510

6472

8278

12675

11406

70

11093

8677

6416

2756

1953

1999

2119

1958

4076

8917

9459

10928

71

10273

8111

7672

2161

1551

1834

1611

1817

5633

9212

8888

9528

72

9121

6247

3640

1903

1568

1453

1062

999

3641

5388

9010

7900

73

6447

5111

3594

2053

2084

1765

1534

1169

3650

5194

8886

8667

74

5965

3672

3206

2120

1522

1297

1208

1864

5269

6917

8598

7939

75

5397

3354

3191

2057

1911

1630

2026

1413

4700

5165

6646

10002

76

8543

6581

4748

2091

1736

1949

1656

2188

5025

8154

10570

12990

77

10970

8114

5215

2676

2406

2292

2388

1644

4737

6157

8854

10408

78

9245

7304

5579

3442

2504

2438

2383

980

3895

7308

8808

12356

79

12614

8856

5592

3616

2865

2779

2549

1207

5086

7406

9276

11719

80

7580

5597

4972

4164

2865

2252

2402

851

4712

8102

9239

13151

81

12457

7682

6931

4389

3353

2698

2269

1128

5946

7667

9178

13151

82

12887

7401

5370

3445

2790

2344

2194

3305

5501

5523

7322

9943

83

9600

7768

7062

5011

3700

3276

2532

1940

4394

6041

9239

11677

84

9488

7432

5762

3115

3177

2803

2446

3697

6040

7767

10120

11852

85

10558

8234

6901

4582

3696

3544

3251

3408

5239

6287

9468

12986

86

11311

7296

6424

5435

3952

3703

3359

2641

6719

7726

9907

10190

AVG

11822

10686

9139

5364

4529

2921

2117

2396

5173

7242

9458

10950


Table 5. Months and areas of maximum catch and effort for northern bluefin tuna caught by Japanese longliners.

YEAR

MO.

AREA

CATCH

MO.

AREA

HOOKS

1952

AUG

140 E 35 N

4571

FEB

140 E 30 N

3010309

1953

MAY

125 E 20 N

4332

FEB

140 E 30 N

5712033

1954

MAY

130 E 20 N

2777

MAR

140 E 30 N

6754806

1955

AUG

140 E 40 N

4695

FEB

140 E 30 N

4918182

1956

AUG

140 E 40 N

8785

JAN

160 E 30 N

2859595

1957

AUG

140 E 35 N

1495

MAR

140 E 30 N

2896375

1958

SEP

170 E 40 N

517

FEB

140 E 30 N

3244643

1959

APR

135 E 30 N

3707

MAR

135 E 30 N

3877797

1960

APR

135 E 30 N

14505

APR

135 E 30 N

3983831

1961

APR

135 E 30 N

6612

APR

135 E 30 N

4178016

1962

APR

135 E 25 N

5491

APR

135 E 30 N

2897219

1963

MAY

125 E 20 N

12733

MAR

135 E 30 N

4882053

1964

MAY

125 E 20 N

2970

APR

135 E 30 N

2892769

1965

MAY

125 E 20 N

2102

FEB

140 E 30 N

1996443

1966

MAY

125 E 25 N

798

OCT

155 E 35 N

4002019

1967

MAY

130 E 25 N

470

OCT

150 E 35 N

4394869

1968

MAR

170 E 30 N

571

OCT

155 E 35 N

2924598

1969

MAY

125 E 25 N

168

NOV

150 E 30 N

2486019

1970

JUNE

135 E 30 N

184

NOV

150 E 30 N

2250384

1971

MAR

170 E 30 N

310

MAR

170 E 45 S

3224520

1972

DEC

170 E 30 N

73

MAR

170 E 45 S

3273213

1973

JUNE

175 E 35 S

174

JULY

150 E 35 S

2034063

1974

JUNE

175 E 35 S

84

FEB

170 E 45 S

1914188

1975

SEP

140 E 45 N

405

MAR

170 E 45 S

2596276

1976

AUG

140 E 40 N

182

MAR

170 E 45 S

3336438

1977

APR

135 E 30 N

703

MAR

170 E 45 S

3920323

1978

SEP

155 E 40 N

1230

OCT

160 E 35 N

2084425

1979

SEP

155 E 40 N

509

MAR

170 E 45 S

3835096

1980

APR

135 E 30 N

88

MAR

170 E 45 S

4460543

1981

MAR

140 E 35 N

365

APR

170 E 45 S

3947549

1982

JULY

145 E 30 N

408

JULY

175 E 35 S

3220419

1983

MAR

140 E 30 N

167

MAY

80 W 0 N

2896969

1984

JAN

140 E 30 N

132

APR

170 E 45 S

2396459

1985

MAR

140 E 35 N

34

DEC

140 E 30 N

2526485

1986

JAN

140 E 30 N

36

DEC

140 E 30 N

2545577


Table 6. Number of 5-degree-month strata, within the core area, with Japanese longline catches of northern bluefin tuna.

YEAR

JAN.

FEB.

MAR.

APR

MAY

JUNE

JULY

AUG.

SEP.

OCT.

NOV.

DEC.

1952

9

9

5

10

17

10

5

6

4

8

11

9

1953

11

18

4

12

16

18

7

8

5

7

13

9

1954

10

17

9

15

17

20

3

4

8

15

14

9

1955

14

13

13

12

18

15

5

12

14

10

10

9

1956

10

12

9

15

14

11

6

5

6

10

5

7

1957

12

16

12

19

10

8

4

3

8

7

7

13

1958

13

16

13

14

7

8

1

5

9

11

11

15

1959

13

17

19

17

11

14

7

14

12

16

15

11

1960

18

20

24

21

19

18

10

9

14

14

11

13

1961

11

14

24

19

19

18

12

13

15

11

12

11

1962

13

21

23

21

16

16

12

9

12

16

13

15

1963

16

19

20

21

13

12

6

7

12

10

11

20

1964

15

13

13

16

16

17

13

8

9

11

7

10

1965

11

22

19

21

23

21

14

11

13

12

16

11

1966

9

14

19

20

17

12

11

12

14

12

18

14

1967

17

19

23

22

15

9

6

11

12

9

9

12

1968

7

13

15

14

12

8

4

8

10

11

11

6

1969

9

10

13

15

8

7

4

9

12

5

8

12

1970

14

12

11

6

8

10

4

3

8

6

10

10

1971

9

12

13

6

8

5

1

1

7

9

6

9

1972

10

7

4

7

5

3

2

1

3

3

5

10

1973

7

5

8

5

5

6

2

3

4

3

11

9

1974

4

7

5

6

6

3

2

7

6

5

10

10

1975

5

3

4

7

7

4

1

4

3

2

2

5

1976

7

5

8

5

6

4

3

3

2

6

8

10

1977

15

14

7

9

5

4

3

1

7

7

2

11

1978

13

9

4

7

7

4

1

3

8

4

4

14

1979

16

15

7

9

12

11

1

4

7

9

8

14

1980

11

9

10

9

8

5

3

0

5

3

12

14

1981

19

18

12

14

7

8

1

5

8

11

19

18

1982

22

16

11

11

10

10

7

5

9

11

7

12

1983

15

9

12

8

7

3

3

4

4

6

11

8

1984

11

13

8

5

2

3

3

5

4

6

11

11

1985

14

11

7

8

4

4

2

1

3

4

9

9

1986

8

11

9

6

1

4

0

2

5

3

7

5

AVG

12

13

12

12

11

10

5

6

8

8

10

11


Table 7. Number of 5-degree-month strata with Japanese longline effort within the core area.

YEAR

JAN.

FEB.

MAR.

APR

MAY

JUNE

JULY

AUG.

SEP.

OCT.

NOV.

DEC.

1952

26

27

28

25

25

16

11

13

16

19

26

27

1953

27

27

24

18

24

27

22

15

14

25

32

27

1954

29

26

27

30

25

25

20

15

22

32

33

33

1955

37

38

36

33

26

25

19

22

30

37

40

39

1956

37

35

33

30

30

24

18

16

29

41

42

39

1957

39

37

29

26

23

19

12

20

28

33

38

35

1958

35

31

27

29

22

21

13

24

24

30

32

35

1959

34

27

29

27

24

22

21

23

38

41

39

34

1960

34

36

35

30

28

33

22

21

29

39

39

37

1961

40

31

33

30

26

30

27

25

25

35

39

42

1962

37

33

41

34

23

26

22

17

27

42

47

41

1963

36

37

37

29

22

19

21

23

31

41

44

49

1964

39

34

32

26

26

25

30

21

33

43

48

44

1965

35

40

40

30

32

35

33

37

38

42

51

43

1966

43

34

32

29

30

31

27

28

37

47

42

41

1967

38

36

31

33

27

28

31

31

39

43

44

46

1968

40

35

33

26

21

23

28

26

35

35

50

45

1969

41

34

31

26

23

19

31

22

42

35

43

45

1970

42

34

26

22

19

26

31

26

35

41

40

42

1971

43

39

41

21

18

27

24

20

32

38

38

39

1972

39

37

31

26

19

18

21

20

20

26

36

41

1973

37

33

34

25

22

27

22

19

26

28

42

38

1974

35

34

26

21

20

19

21

18

23

29

38

35

1975

37

30

30

24

22

21

18

14

19

20

31

36

1976

34

36

38

25

26

26

27

26

32

43

46

42

1977

35

32

34

28

19

21

29

22

31

37

38

38

1978

33

29

29

21

27

22

21

11

32

30

35

35

1979

29

26

25

24

28

26

26

23

30

36

37

38

1980

34

27

27

24

23

20

16

16

27

35

41

42

1981

35

35

35

30

30

30

27

24

36

44

48

40

1982

35

35

31

29

24

21

19

22

28

33

35

39

1983

33

32

25

26

26

23

24

22

32

33

40

37

1984

35

34

29

24

23

25

20

25

36

32

34

36

1985

35

33

34

24

17

14

18

23

25

28

37

38

1986

33

36

33

26

14

15

21

25

31

32

36

36

AVG

36

33

32

27

24

24

23

22

29

35

39

39


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