Table 26: Standard length (Sl, mm) of Carassius auratus from the Anzali Lagoon sampled in 1991. Rosa Lee method, correction factor 22.6 mm.
Age group | n | Sl at capture | Back calculated Sl for the previous growth seasons | |||||
l1 | l2 | l3 | l4 | l5 | l6 | |||
I | 2 | 135 | 100 | |||||
II | 7 | 175 | 88 | 133 | ||||
III | 24 | 219 | 89 | 142 | 183 | |||
IV | 24 | 258 | 95 | 148 | 191 | 227 | ||
V | 9 | 275 | 87 | 134 | 183 | 222 | 252 | |
VI | 1 | 320 | 99 | 145 | 201 | 247 | 268 | 297 |
Total | 67 | Grand average | 93 | 140 | 190 | 232 | 260 | 297 |
Increments in mm | 93 | 47 | 50 | 42 | 28 | 37 | ||
Increments in % of the l1 length | 100 | 50 | 54 | 45 | 30 | 40 | ||
C1h | 38.0 | 42.8 | 37.9 | 26.4 | 34.6 |
Table 27: Linear growth of Carassius auratus in various waterbodies outside Iran. Only the best growing populations are considered.
Locality | Linf. | K | to | Source |
Kartal Lake | 373.2 | 0.3982 | 0.0161 | 1 |
Trstená arm (middle Danube) | 293.0 | 0.3385 | 0.0239 | 2 |
Danube delta USSR | 398.9 | 0.2829 | 0.0495 | 1 |
Anzali Lagoon | 385.4 | 0.2389 | 0.0654 | 3 |
Danube delta (Roumania) | 429.3 | 0.2091 | -0.0094 | 4 |
Chervonoe Lake (Belorussia) | 446.0 | 0.1991 | -0.0094 | 5 |
Kolárovo canal (Danube basin, Slovakia) | 539.9 | 0.1733 | 0.0109 | 6 |
Polkovnikovo Lake (northern Kazakhstan) | 486.8 | 0.1507 | 0.0667 | 7 |
Note: The Von Bertalanffy equation in this and other tables of this type were calculated from data of other authors:
1 - Kukuradze and Mariyash (1975),
2 - Černý (1990),
3 - our data,
4 - Papadopol and Mester (1980),
5 - Zhukov (1965),
6 - Makara (1979),
7 - Ereshchenko (1959)
Table 28: Linear growth of Cyprinus carpio from various localities of the southern Caspian Sea and coastal waters.
Locality | Linf. | K | to | source |
Astara River 1926 | 529.0 | 0.3037 | -0.0446 | 1 |
Kura River 1926 | 700.3 | 0.2531 | -0.0353 | 2 |
Sarysu Lake 1933 | 770.8 | 0.2510 | 0.2479 | 2 |
Southern Caspian 1933 | 946.3 | 0.1694 | -0.0135 | 1 |
Anzali Lagoon 1991 | 777.3 | 0.1536 | 0.0842 | 3 |
Soyukhbulag 1960 | 1653.5 | 0.0529 | 0.0133 | 2 |
1 - Nikol'skii 1949;
2 - Abdurakhmanov (1962);
3 - our data
Table 29: Catch of Rutilus frisii kutum in the Iranian waters of the Caspian Sea in 1987–1991.
Year | Catch (t) |
1987 | 3500 |
1988 | 3500*) |
1989 | 5000*) |
1990 | 6500 |
1991 | 8855 |
Table 30: Number of Rutilus frisii kutum fingerlings (million) released into Iranian waters of the Caspian Sea watershed in 1986–1991
Years | Total number released | Number released in the Anzali area ( % in brackets) | |
1985 | 38 | ? | ? |
1986 | 52 | 15 | (29) |
1987 | 72 | 18 | (25) |
1988 | 84 | 20 | (24) |
1989 | 140 | 30 | (21) |
1990 | 142 | 35 | (25) |
1991 | 170 | 35 | (21) |
Table 31: Differences in length (Sl, Slj1, Slj2; mm) in the young-of-the-year juveniles of Rutilus frisii kutum caught in Sowsar roga on 14.4.1991. Based on a number of juvenile marks present on scales. t = test between juveniles with one and two juvenile marks. **=P< 0.05. Mean (above) and ranges (below).
Number of juvenile marks | n | Sl at capture | Calculated Sl at formation of juvenile marks | |
Slj1 | Slj2 | |||
0 | 2 | 65.5 | ||
58 – 73 | ||||
1 | 26 | 66.7 | 39.6 | |
53 – 82 | 27 – 50 | |||
2 | 10 | 68.8 | 33.1 | 47.1 |
61 – 81 | 25 – 46 | 30 – 55 | ||
t | 0.74 | 3.14** |
Table 32: Age composition by percent of Rutilus frisii kutum (numerator - one year, denominator - two years).
Age group | I | II | III | IV | V | VI | VII | VIII |
Males | - | - | 12 | 35 | 47 | 6 | - | - |
- | - | - | 11 | 56 | 22 | 9 | 2 | |
Females | - | - | - | 20 | 40 | 40 | - | - |
- | - | - | 13 | 27 | 40 | 13 | 7 |
Table 33: Age/length key for Rutilus frisii kutum sample taken in the Anzali Lagoon in 1991. Data are percentage per 50 mm length groups
Years spent in freshwater after hatching | ||||||||||||||||
1 | 2 | |||||||||||||||
age group (year) | length group (mm) | age group (year) | ||||||||||||||
VIII | VII | VI | V | IV | III | II | I | I | II | III | IV | V | VI | VII | VIII | |
- | - | - | - | - | - | - | 100 | 51 – 100 | - | 100 | - | - | - | - | - | - |
- | - | - | - | - | - | - | - | 101 – 150 | - | - | - | - | - | - | - | - |
- | - | - | - | - | - | - | - | 151 – 200 | - | - | - | - | - | - | - | - |
- | - | - | - | - | - | - | - | 201 – 250 | - | - | - | - | - | - | - | - |
- | - | - | - | - | 100 | - | - | 251 – 300 | - | - | - | 100 | - | - | - | - |
- | - | - | 45 | 33 | 22 | - | - | 301 – 350 | - | - | - | 22 | 56 | 22 | - | - |
- | - | 5 | 60 | 30 | 5 | - | - | 351 – 400 | - | - | - | 4 | 69 | 23 | 4 | - |
- | - | 30 | 70 | - | - | - | - | 401 – 450 | - | - | - | - | 27 | 46 | 27 | - |
- | - | 100 | - | - | - | - | - | 451 – 500 | - | - | - | - | - | 33 | 33 | 33 |
- | - | 100 | - | - | - | - | - | 501 – 550 | - | - | - | - | - | 50 | 50 | - |
- | - | - | - | - | - | - | - | 551 – 600 | - | - | - | - | - | - | - | 100 |
Table 34: Growth in length (Sl, mm) of Rutilus frisii kutum males calculated from a sample taken in the Anzali Lagoon in 1991, separated according to number of years spent in freshwater after hatching (above-one year, below-two years). Absolute (in mm) and relative (in %) increments are also shown, as well as coefficient of condition (K), and weight at capture (w in grams).
Age group | n | Size at capture | Back calculated Sl for the previous growth seasons | ||||||||||
Sl | w | K | l0 | l1 | l2 | l3 | l4 | l5 | l6 | l7 | l8 | ||
I | 38* | 67 | - | - | - | 67* | |||||||
II | 0 | - | - | - | - | - | - | ||||||
III | 3 | 77 | - | - | 56 | 56 | 77 | ||||||
4 | 334 | 467 | 1.20 | 44 | 127 | 230 | 334 | ||||||
IV | - | - | - | - | - | - | - | - | |||||
12 | 344 | 654 | 1.58 | 44 | 77 | 180 | 281 | 344 | |||||
V | 5 | 310 | 575 | 1.68 | 36 | 65 | 99 | 204 | 310 | ||||
16 | 377 | 800 | 1.55 | 38 | 62 | 140 | 240 | 319 | 377 | ||||
VI | 26 | 370 | 757 | 1.52 | 38 | 58 | 100 | 210 | 308 | 369 | |||
2 | 358 | 500 | 1.21 | 39 | 63 | 123 | 188 | 251 | 298 | 358 | |||
VII | 10 | 367 | 819 | 1.53 | 39 | 66 | 100 | 165 | 245 | 321 | 367 | ||
- | - | - | - | - | - | - | - | - | - | - | - | ||
VIII | 4 | 426 | 1283 | 1.56 | 44 | 55 | 89 | 162 | 238 | 320 | 393 | 426 | |
- | - | - | - | - | - | - | - | - | - | - | - | - | |
1 | 460 | 1400 | 1.44 | 43 | 74 | 118 | 161 | 241 | 281 | 381 | 422 | 460 | |
Total | 34 | Grand average | 41 | 82 | 168 | 261 | 315 | 338 | 358 | - | - | ||
49 | 40 | 62 | 99 | 180 | 268 | 323 | 380 | 424 | 460 | ||||
Increments in mm | 41 | 82 | 86 | 93 | 54 | 23 | 20 | - | - | ||||
40 | 62 | 37 | 81 | 88 | 55 | 57 | 44 | 36 | |||||
Increments in % of | 100 | 105 | 113 | 66 | 28 | 24 | - | - | |||||
100 | 60 | 131 | 142 | 89 | 92 | 71 | 58 |
* Not included in the total sum and grand everage respectively
Table 35: Growth in length of Rutilus frisii kutum females from the same sample as for males. Explanation of symbols same as in Table 34.
Age group | n | Size at capture | Back calculated Sl for the previous growth seasons | ||||||||||
Sl | w | K | l0 | l1 | l2 | l3 | l4 | l5 | l6 | l7 | l8 | ||
IV | 4 | 388 | 760 | 1.32 | 48 | 71 | 184 | 304 | 388 | ||||
1 | 325 | 500 | 1.46 | - | 67 | 106 | 221 | 325 | |||||
V | 5 | 423 | 1317 | 1.65 | 43 | 90 | 177 | 258 | 367 | 423 | |||
5 | 415 | 1100 | 1.61 | 45 | 69 | 115 | 258 | 361 | 415 | ||||
VI | 6 | 463 | 1680 | 1.83 | 36 | 57 | 140 | 247 | 359 | 420 | 463 | ||
6 | 451 | 1360 | 1.64 | 41 | 57 | 96 | 186 | 308 | 399 | 447 | |||
VII | - | - | - | - | - | - | - | - | - | - | - | - | |
2 | 481 | - | - | 31 | 44 | 70 | 162 | 230 | 283 | 364 | 472 | ||
VIII | - | - | - | - | - | - | - | - | - | - | - | - | - |
1 | 560 | - | - | - | 58 | 95 | 154 | 213 | 261 | 362 | 424 | 517 | |
Total | 16 | Grand average | 42 | 73 | 167 | 270 | 371 | 422 | 463 | - | - | ||
15 | 39 | 59 | 96 | 196 | 287 | 340 | 391 | 448 | 517 | ||||
Increments in mm | 31 | 94 | 103 | 101 | 103 | 51 | 41 | - | - | ||||
20 | 59 | 37 | 100 | 91 | 53 | 51 | 57 | 69 | |||||
Increments in % of the l1 length | 100 | 110 | 107 | 110 | 54 | 43 | - | - | |||||
100 | 63 | 169 | 154 | 90 | 86 | 97 | 117 |
Table 36: Parameters of the von Bertalanffy equation characterizing linear growth of Rutilus frisii kutum from various localities
Locality | Linf. | K | to | source |
males | ||||
Anzali Lagoon 1969–1970 | 706.2 | 0.12374 | -0.0261 | 1 |
Anzali Lagoon 1991 (1 yr spent in freshwater) | 487.4 | 0.2259 | -0.0895 | 2 |
Malyi Kyzylagach Bay 1960 | 790.8 | 0.1853 | 0.0627 | 3 |
Anzali Lagoon 1991 (2 yrs spent in freshwater) | 1360.4 | 0.0516 | -0.2083 | 2 |
females | ||||
Anzali Lagoon 1961–1970 | 804.2 | 0.2070 | -0.0006 | 1 |
Malyi Kyzylagach Bay 1960 | 805.6 | 0.1826 | 0.0347 | 3 |
Anzali Lagoon 1914–1915 | 811.8 | 0.1607 | -0.0752 | 4 |
Western Bank of Middle Caspian | 1002.0 | 0.1061 | -0.0436 | 4 |
Anzali Lagoon 1991 (1 yr spent in freshwater) | 1042.0 | 0.0982 | -0.0542 | 2 |
Bay of Astrabad | 1137.0 | 0.0870 | -0.0114 | 4 |
Bay of Kura | 1302.3 | 0.0823 | -0.0198 | 4 |
Anzali Lagoon 1991 (2 yrs spent in freshwater) | 6202.7 | 0.0109 | 0.1840 | 2 |
1 - RaLonde and Razavi (1972),
2 - our data,
3 - Abdurakhmanov,
4 - Berg (1948–1949)
Table 37: Growth characteristics of the Rutilus frisii kutum in the Anzali Lagoon in 1970 and 1990. Figures from 1969–1970 calculated from data by RaLonde and Razavi (1972)
Sample analyzed (years) | C 1h between years of life | ||||||||||||||
1 | 2 | 3 | 4 | 5 | 6 | 7 | 8 | ||||||||
Males | |||||||||||||||
1969–1970 | 55.2 | 45.9 | 25.7 | 18.9 | |||||||||||
1990–19911) | 58.6 | 74.0 | 49.1 | 22.2 | 19.4 | ||||||||||
1990–19912) | 29.0 | 59.2 | 71.6 | 50.0 | 52.5 | 41.6 | 34.6 | ||||||||
Females | |||||||||||||||
1969–1970 | 64.7 | 41.3 | 26.7 | 24.5 | 21.0 | ||||||||||
1990–19911) | 60.4 | 80.2 | 85.8 | 47.8 | 39.2 | ||||||||||
1990–19912) | 28.7 | 68.5 | 74.7 | 48.6 | 47.5 | 53.2 | 64.2 |
1) - one year spent in freshwaters;
2) - two years spent in freshwaters
Table 38: Growth in length (Sl, mm) of Chalcalburnus chalcoides from the Anzali Lagoon in 1991
Age group | n | Sl at capture | Back calculated Sl for the previous growth seasons | |||||
l1 | l2 | l3 | l4 | l5 | ||||
II | 6 | 111 | 81 | 111 | ||||
III | 50 | 129 | 73 | 107 | 129 | |||
IV | 21 | 173 | 78 | 113 | 142 | 173 | ||
V | 3 | 189 | 73 | 108 | 130 | 157 | 189 | |
Total | 80 | Grand average | 76 | 110 | 134 | 165 | 189 | |
Increments in mm | 76 | 34 | 24 | 31 | 24 | |||
Increments in % of the l1 length | 100 | 45 | 32 | 41 | 32 | |||
Clh | 28.1 | 21.7 | 27.9 | 22.4 |
Table 39: Parameters of the von Bertalanffy equation characterizing linear growth of Chalcalburnus chalcoides from various localities in the Caspian Sea basin. Values other than for Anzali Lagoon calculated from data introduced by Abdurakhmanov (1962)
Locality | Linf. | K | to |
Kura (Shikhovo) 1945–1950 | 271.2 | 0.6101 | -0.0271 |
Kura 1930's | 358.8 | 0.4206 | 0.0007 |
Kura (Bank) 1948–1949 | 342.9 | 0.4135 | 0.0128 |
Kura (Bank) 1949–1950 | 343.0 | 0.4170 | -0.0134 |
Kura (Pirsagat) 1940–1950 | 362.6 | 0.4088 | -0.0149 |
Anzali Lagoon 1991 | 218.2 | 0.3739 | 0.0557 |
Kura 1966–1968 | 387.7 | 0.3558 | 0.0242 |
Kura (Khudat) 1930 | 405.6 | 0.2977 | 0.0460 |
Lenkoran 1954 | 362.3 | 0.2400 | 0.0031 |
Zyuidostav Kultuk 1945–1950 | 412.8 | 0.2054 | 0.0049 |
Table 40: Growth in length (Sl, mm) of Vimba vimba from the Anzali Lagoon in 1991
Age group | n | Sl at capture | Back calculated Sl for the previous growth seasons | |||||
l1 | l2 | l3 | l4 | l5 | l6 | |||
III | 48 | 145 | 70 | 109 | 145 | |||
IV | 70 | 171 | 67 | 104 | 138 | 171 | ||
V | 20 | 200 | 64 | 102 | 141 | 173 | 200 | |
VI | 2 | 225 | 55 | 96 | 130 | 169 | 198 | 225 |
Total | 140 | Grand average | 64 | 103 | 139 | 171 | 199 | 225 |
Increments in mm | 64 | 39 | 36 | 32 | 28 | 26 | ||
Increments in % of the l1 length | 100 | 61 | 56 | 50 | 44 | 41 | ||
Clh | 30.4 | 30.9 | 28.8 | 25.9 | 24.4 |
Table 41: Parameters of the von Bertalanffy equation characterizing linear growth of Vimba vimba from various localities in the Caspian Sea basin. Values other than for Anzali lagoon calculated from data introduced by Abdurakhmanov (1962)
Locality | Linf. | K | to |
Terek River (1940's ?) | 284.0 | 0.3419 | 0.0625 |
Dagestan 1910 | 296.6 | 0.3180 | 0.0639 |
Anzali Lagoon 1991 | 324.1 | 0.1952 | 0.0425 |
Terek River 1952 | 331.5 | 0.1783 | 0.0553 |
Sara Lake 1950's | 356.6 | 0.1323 | -0.0279 |
Table 42: Length composition, total mortality rate (A), survival rate (S) and instantaneous rate of total mortality (2) of Astacus leptodactulus in the Anzali Lagoon in 1990
Size group (OCL, mm)
Region | 20–24 | 25–29 | 30–34 | 35–39 | 40–44 | 45–49 | 50–54 | Σ | |
n | - | 118 | 292 | 133 | 40 | 9 | - | 592 | |
% | - | 19.9 | 49.3 | 22.5 | 6.8 | 1.5 | - | 100 | |
Seyjan | A | - | - | 0.5445 | 0.6933 | 0.775 | - | - | 0.6086 |
S | - | - | 0.4555 | 0.3007 | 0.2250 | - | - | 0.3914 | |
Z | - | - | 0.78 | 1.20 | 1.49 | - | - | 0.94 | |
n | 1 | 103 | 445 | 360 | 281 | 95 | 1 | 1.286 | |
% | 0.1 | 8.0 | 34.6 | 28.0 | 21.8 | 7.4 | 0.1 | 100 | |
Siahdar -vishan | A | - | - | 0.1910 | 0.2194 | 0.6619 | 0.9895 | - | 0.3760 |
S | - | - | 0.8090 | 0.7806 | 0.3381 | 0.0105 | - | 0.6240 | |
Z | - | - | 0.21 | 0.25 | 1.08 | 4.56 | - | 0.47 |